The speckled wood can be seen in dappled sunlight in woodlands. The male tends to perch in patches of sunlight, and intercepts intruding butterflies. They may also patrol an area in search of females. This species does not usually feed on flowers but males and females feed on honeydew produced by aphids up in the tree canopy. There are three or more generations per year, Females lay their eggs singly on grass. The caterpillars, which hatch after around ten days, are very well camouflaged against the blades of grass, thanks to their green colouration. Pupae form attached to the foodplant or to vegetation nearby. The speckled wood can overwinter either as a caterpillar or as a pupa.